Dick Dastardly and Muttley, the comic villains from Wacky Races, are flying aces and members of the Vulture Squadron, a crew of aviators on a mission to stop a homing pigeon named Yankee Doodle Pigeon from delivering messages to the other side.
Each story features variations on the same plot elements: the Vulture Squadron tries to trap Yankee Doodle Pigeon using one or more planes equipped with Klunk's latest contraptions, but one or more of the Squadron messes up and the plane(s) either crash, collide or explode (or even all of the above!). While they are falling out of the wreckage, Dick Dastardly calls for help, which Muttley either offers or refuses depending on whether Dastardly agrees to give him a medal. Even when Muttley does agree to fly Dastardly out of trouble, Dastardly seldom has a soft landing. At some point the General calls Dastardly on the phone to demand results. Dastardly assures him that they will soon capture the pigeon.